@font-face{font-family:Editorial NBSP;src:local(Iowan Old Style Roman),local(IowanOldStyle-Roman),local(Cambria),local(Georgia);unicode-range:U+A0}:root{color-scheme:light;--font-ui:-apple-system, BlinkMacSystemFont, "Segoe UI", Roboto, Helvetica, Arial, sans-serif;--font-editorial:"Editorial NBSP", Charter, "Bitstream Charter", "Sitka Text", Cambria, Georgia, serif;--font-mono:ui-monospace, SFMono-Regular, "SF Mono", Menlo, Consolas, monospace;--text-caption:.75rem;--text-meta:.8125rem;--text-small:.875rem;--text-body:1rem;--ink:#23211c;--ink-soft:#383429;--muted:#67614f;--paper:#f5f2e9;--paper-deep:#e8e2d2;--paper-bright:#fcfaf5;--amber:#e3b04b;--amber-soft:#e3b04b38;--amber-deep:#b8842a;--amber-text:#7c5510;--sea:#1d6b78;--sea-soft:#1d6b781f;--sea-text:#145761;--sea-bright:#2b8a99;--rust:#ab5330;--rust-dark:#7d3d24;--on-rust:#fff;--line:#23211c29;--line-strong:#23211c4d;--control-border:#8a8474;--control-bg:#fff;--surface-glass:#f8f4eae6;--shadow:0 28px 70px #23211c21;--shadow-soft:0 10px 30px #23211c14;--inverse-bg:#23211c;--inverse-text:#fcfaf5;--inverse-muted:#fcfaf5b8;--on-accent:#23211c;--success:#3d5f3a;--search-icon:url("data:image/svg+xml,%3Csvg xmlns='http://www.w3.org/2000/svg' width='21' height='21' fill='none' stroke='%2367614f' stroke-width='2' viewBox='0 0 24 24'%3E%3Ccircle cx='11' cy='11' r='7'/%3E%3Cpath d='m20 20-4-4'/%3E%3C/svg%3E");font-family:var(--font-ui);font-synthesis:none;text-rendering:optimizelegibility}:root[data-theme=dark]{color-scheme:dark;--ink:#f0ede2;--ink-soft:#d5d0c1;--muted:#aca695;--paper:#14161a;--paper-deep:#1a1d22;--paper-bright:#22262c;--amber:#e3b04b;--amber-soft:#e3b04b24;--amber-deep:#d29f3c;--amber-text:#e8bd66;--sea:#6cbecb;--sea-soft:#6cbecb24;--sea-text:#7fcbd7;--sea-bright:#8ad4e0;--rust:#d5825b;--rust-dark:#eda27c;--on-rust:#14161a;--line:#f0ede224;--line-strong:#f0ede245;--control-border:#7f8894;--control-bg:#1e2228;--surface-glass:#1a1d22eb;--shadow:0 28px 70px #0000006b;--shadow-soft:0 10px 30px #00000047;--inverse-bg:#0d0f12;--inverse-text:#f6f4ec;--inverse-muted:#f6f4ecb3;--on-accent:#14161a;--success:#92b98c;--search-icon:url("data:image/svg+xml,%3Csvg xmlns='http://www.w3.org/2000/svg' width='21' height='21' fill='none' stroke='%23aca695' stroke-width='2' viewBox='0 0 24 24'%3E%3Ccircle cx='11' cy='11' r='7'/%3E%3Cpath d='m20 20-4-4'/%3E%3C/svg%3E")}@media (prefers-color-scheme:dark){:root:not([data-theme]){color-scheme:dark;--ink:#f0ede2;--ink-soft:#d5d0c1;--muted:#aca695;--paper:#14161a;--paper-deep:#1a1d22;--paper-bright:#22262c;--amber-soft:#e3b04b24;--amber-deep:#d29f3c;--amber-text:#e8bd66;--sea:#6cbecb;--sea-soft:#6cbecb24;--sea-text:#7fcbd7;--sea-bright:#8ad4e0;--rust:#d5825b;--rust-dark:#eda27c;--on-rust:#14161a;--line:#f0ede224;--line-strong:#f0ede245;--control-border:#7f8894;--control-bg:#1e2228;--surface-glass:#1a1d22eb;--shadow:0 28px 70px #0000006b;--shadow-soft:0 10px 30px #00000047;--inverse-bg:#0d0f12;--inverse-text:#f6f4ec;--inverse-muted:#f6f4ecb3;--on-accent:#14161a;--success:#92b98c;--search-icon:url("data:image/svg+xml,%3Csvg xmlns='http://www.w3.org/2000/svg' width='21' height='21' fill='none' stroke='%23aca695' stroke-width='2' viewBox='0 0 24 24'%3E%3Ccircle cx='11' cy='11' r='7'/%3E%3Cpath d='m20 20-4-4'/%3E%3C/svg%3E")}}*{box-sizing:border-box}html{scroll-behavior:smooth}body{background:radial-gradient(circle at 84% 6%, var(--sea-soft), transparent 26rem), radial-gradient(circle at 6% 62%, var(--amber-soft), transparent 30rem), var(--paper);min-width:0;color:var(--ink);margin:0;font-size:1rem;line-height:1.55}body:before{z-index:-1;pointer-events:none;content:"";background-image:url("data:image/svg+xml,%3Csvg width='180' height='180' viewBox='0 0 180 180' xmlns='http://www.w3.org/2000/svg'%3E%3Cfilter id='n'%3E%3CfeTurbulence type='fractalNoise' baseFrequency='.8' numOctaves='3' stitchTiles='stitch'/%3E%3C/filter%3E%3Crect width='100%25' height='100%25' filter='url(%23n)' opacity='.028'/%3E%3C/svg%3E");position:fixed;inset:0}main{overflow:clip}a{color:inherit}img,svg{max-width:100%}img{display:block}button,input,select,textarea{font:inherit}[hidden]{display:none!important}::selection{background:var(--amber);color:var(--on-accent)}:focus-visible{outline:3px solid var(--sea);outline-offset:3px}.visually-hidden{clip:rect(0, 0, 0, 0)!important;white-space:nowrap!important;border:0!important;width:1px!important;height:1px!important;margin:-1px!important;padding:0!important;position:absolute!important;overflow:hidden!important}.shell{width:min(1180px,100% - 48px);margin-inline:auto}.shell-narrow{width:min(760px,100% - 48px);margin-inline:auto}.skip-link{z-index:100;background:var(--inverse-bg);color:var(--inverse-text);border-radius:999px;padding:12px 18px;font-weight:750;text-decoration:none;position:fixed;top:12px;left:12px;transform:translateY(-180%)}.skip-link:focus{transform:translateY(0)}.site-header{z-index:20;border-bottom:1px solid var(--line);justify-content:space-between;align-items:center;gap:16px;min-height:86px;display:flex;position:relative}.brand{letter-spacing:-.025em;align-items:center;gap:10px;font-size:1.05rem;font-weight:850;text-decoration:none;display:inline-flex}.brand svg{width:27px;height:auto;color:var(--ink);flex:none}.brand-wordmark{white-space:nowrap}.brand-wordmark>span{color:var(--sea-text);font-weight:720}.header-actions{align-items:center;gap:12px;display:flex}.desktop-nav{align-items:center;gap:clamp(10px,1.7vw,24px);display:flex}.desktop-nav a{color:var(--muted);padding-block:32px;font-size:.84rem;font-weight:760;text-decoration:none;position:relative}.desktop-nav a:after{background:var(--rust);content:"";opacity:0;height:3px;transition:opacity .16s;position:absolute;bottom:-1px;left:0;right:0}.desktop-nav a:hover,.desktop-nav a[aria-current]{color:var(--ink)}.desktop-nav a[aria-current]:after{opacity:1}.header-search-link,.theme-picker>summary,.mobile-nav>summary{border:1px solid var(--line-strong);cursor:pointer;width:40px;height:40px;color:var(--ink);background:0 0;border-radius:999px;place-items:center;list-style:none;display:grid}.header-search-link::marker,.theme-picker>summary::marker,.mobile-nav>summary::marker{display:none}.header-search-link::-webkit-details-marker{display:none}.theme-picker>summary::-webkit-details-marker{display:none}.mobile-nav>summary::-webkit-details-marker{display:none}.header-search-link{text-decoration:none}.header-search-link span{background:var(--search-icon) center / contain no-repeat;width:21px;height:21px}.theme-picker{position:relative}.theme-picker>summary svg{width:19px;height:19px}.theme-picker-menu{z-index:30;border:1px solid var(--line-strong);background:var(--surface-glass);min-width:176px;box-shadow:var(--shadow-soft);backdrop-filter:blur(10px);border-radius:16px;padding:8px;display:grid;position:absolute;top:calc(100% + 10px);right:0}.theme-picker-menu button{color:var(--ink);cursor:pointer;font-size:var(--text-small);text-align:left;background:0 0;border:0;border-radius:10px;justify-content:space-between;align-items:center;gap:12px;padding:9px 12px;font-weight:650;display:flex}.theme-picker-menu button:hover{background:var(--paper-deep)}.theme-picker-menu button[aria-pressed=true]:after{color:var(--sea-text);content:"✓";font-weight:800}.mobile-nav{display:none;position:relative}.mobile-nav>summary span{background:var(--ink);border-radius:2px;width:17px;height:2px;margin-block:2px;display:block}.mobile-nav nav{z-index:30;border:1px solid var(--line-strong);background:var(--surface-glass);min-width:232px;box-shadow:var(--shadow-soft);backdrop-filter:blur(10px);border-radius:18px;padding:10px;display:grid;position:absolute;top:calc(100% + 10px);right:0}.mobile-nav nav a{color:var(--ink);font-size:var(--text-small);border-radius:11px;padding:11px 13px;font-weight:680;text-decoration:none}.mobile-nav nav a:hover,.mobile-nav nav a[aria-current]{background:var(--paper-deep)}h1,h2,h3,h4{font-family:var(--font-editorial);letter-spacing:-.02em;text-wrap:balance;font-weight:600;line-height:1.08}h1{margin:0;font-size:clamp(2.5rem,6vw,4.6rem)}h2{margin:0;font-size:clamp(1.85rem,3.4vw,2.9rem)}h3{margin:0;font-size:clamp(1.2rem,1.9vw,1.5rem)}p{text-wrap:pretty;margin:0 0 1.05em}.eyebrow{color:var(--rust);font-family:var(--font-ui);font-size:var(--text-caption);letter-spacing:.16em;text-transform:uppercase;margin:0 0 14px;font-weight:800}.card-label{color:var(--amber-text);font-size:var(--text-caption);letter-spacing:.14em;text-transform:uppercase;margin:0 0 12px;font-weight:800}.kicker{color:var(--muted);font-size:var(--text-meta);letter-spacing:.02em;margin:0 0 8px;font-weight:700}.lede{max-width:60ch;color:var(--ink-soft);font-size:clamp(1.05rem,1.5vw,1.24rem);line-height:1.6}.scientific-name{font-style:italic}.section-heading{margin-bottom:40px}.section-heading p{max-width:56ch;color:var(--muted);margin:0}.split-heading{grid-template-columns:minmax(0,1.25fr) minmax(0,1fr);align-items:end;gap:24px 48px;display:grid}.section-action{margin-top:36px}.button{background:var(--inverse-bg);color:var(--inverse-text);font-size:var(--text-small);cursor:pointer;border:1px solid #0000;border-radius:999px;align-items:center;gap:10px;padding:15px 26px;font-weight:780;text-decoration:none;transition:transform .15s,background 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strong:first-child{color:var(--rust)}.figure{margin:2em 0}.figure figcaption{color:var(--muted);font-size:var(--text-meta);margin-top:14px;line-height:1.55}.figure svg{width:100%;height:auto}.table-scroll{border:1px solid var(--line);background:var(--paper-bright);border-radius:18px;margin:1.8em 0;overflow-x:auto}table{border-collapse:collapse;width:100%;font-size:var(--text-small)}th,td{border-bottom:1px solid var(--line);text-align:left;vertical-align:top;padding:13px 18px}thead th{color:var(--muted);font-size:var(--text-caption);letter-spacing:.1em;text-transform:uppercase;white-space:nowrap;font-weight:800}tbody tr:last-child td{border-bottom:0}.card-grid{grid-template-columns:repeat(auto-fill,minmax(min(100%,290px),1fr));gap:22px;display:grid}.card-grid-2{grid-template-columns:repeat(auto-fill,minmax(min(100%,400px),1fr))}.link-card{border:1px solid var(--line);background:var(--paper-bright);color:inherit;border-radius:22px;flex-direction:column;padding:28px;text-decoration:none;transition:border-color .16s,transform .16s;display:flex}.link-card:hover{border-color:var(--sea);transform:translateY(-2px)}.link-card h3{margin-bottom:10px}.link-card p{color:var(--muted);font-size:var(--text-small);margin-bottom:16px}.link-card b{color:var(--sea-text);font-size:var(--text-meta);margin-top:auto;font-weight:780}.link-card>span:first-child{color:var(--amber-text);font-size:var(--text-caption);letter-spacing:.14em;text-transform:uppercase;margin-bottom:18px;font-weight:800;display:block}.genus-grid{grid-template-columns:repeat(auto-fill,minmax(min(100%,300px),1fr));gap:22px;display:grid}.genus-card{border:1px solid var(--line);background:var(--paper-bright);border-radius:22px;flex-direction:column;transition:border-color .16s,transform .16s;display:flex;overflow:hidden}.genus-card:hover{border-color:var(--sea);transform:translateY(-2px)}.genus-card-figure{background:radial-gradient(circle at 50% 40%, var(--paper-deep), transparent 70%);place-items:center;min-height:168px;padding:26px 24px 18px;display:grid;position:relative}.genus-card-figure svg{width:min(100%,220px);height:auto}.genus-card-figure>span{background:var(--inverse-bg);color:var(--inverse-text);font-size:var(--text-caption);letter-spacing:.04em;border-radius:999px;padding:4px 10px;font-weight:760;position:absolute;top:14px;left:16px}.genus-card-body{flex-direction:column;flex:1;padding:6px 26px 26px;display:flex}.genus-card-body h2,.genus-card-body h3{margin:0 0 4px;font-size:1.42rem}.genus-card-body h2 a,.genus-card-body h3 a{text-decoration:none}.genus-card-body h2 a:hover,.genus-card-body h3 a:hover{text-decoration:underline;text-decoration-color:var(--amber);text-underline-offset:4px;text-decoration-thickness:2px}.genus-card-body>p{color:var(--muted);font-size:var(--text-small);margin-bottom:16px;line-height:1.58}.genus-card-meta{border-top:1px solid var(--line);font-size:var(--text-meta);gap:4px;margin-top:auto;padding-top:16px;display:grid}.genus-card-meta div{justify-content:space-between;gap:10px;display:flex}.genus-card-meta dt,.genus-card-meta span:first-child{color:var(--muted)}.genus-card-meta span:last-child{text-align:right;font-weight:700}.range-bar{width:100%;height:auto;display:block}.range-legend{color:var(--muted);font-size:var(--text-caption);flex-wrap:wrap;gap:10px 18px;margin:14px 0 0;padding:0;list-style:none;display:flex}.range-legend li{align-items:center;gap:7px;display:inline-flex}.range-legend i{border-radius:3px;width:13px;height:13px}.explorer{grid-template-columns:268px minmax(0,1fr);align-items:start;gap:32px;display:grid}.explorer-filters{border:1px solid var(--line);background:var(--paper-bright);border-radius:22px;gap:18px;padding:24px;display:grid;position:sticky;top:22px}.explorer-filters h2{font-size:1.15rem}.filter-group{gap:9px;display:grid}.filter-group>span{color:var(--muted);font-size:var(--text-caption);letter-spacing:.1em;text-transform:uppercase;font-weight:800}.filter-group label{font-size:var(--text-small);align-items:center;gap:9px;display:flex}.filter-group input[type=checkbox],.filter-group input[type=radio]{accent-color:var(--sea);width:16px;height:16px}.search-field{position:relative}.search-field input{border:1px solid var(--control-border);background:var(--control-bg) var(--search-icon) 15px center / 19px no-repeat;width:100%;color:var(--ink);border-radius:999px;padding:13px 16px 13px 44px}.search-field input::placeholder{color:var(--muted)}.explorer-results-head{color:var(--muted);font-size:var(--text-small);flex-wrap:wrap;justify-content:space-between;align-items:baseline;gap:12px;margin-bottom:22px;display:flex}.explorer-empty{border:1px dashed var(--line-strong);color:var(--muted);text-align:center;border-radius:22px;padding:44px 28px}select{border:1px solid var(--control-border);background:var(--control-bg);color:var(--ink);font-size:var(--text-small);border-radius:10px;padding:10px 14px}.profile-hero{grid-template-columns:minmax(0,1.05fr) minmax(0,.95fr);align-items:start;gap:44px;padding-block:clamp(38px,6vw,70px);display:grid}.profile-figure{border:1px solid var(--line);background:radial-gradient(circle at 50% 35%, var(--paper-deep), transparent 72%), var(--paper-bright);border-radius:26px;place-items:center;padding:34px;display:grid}.profile-figure svg{width:min(100%,340px);height:auto}.profile-figure figcaption{color:var(--muted);font-size:var(--text-meta);text-align:center;margin-top:18px}.fact-table{border:1px solid var(--line);background:var(--paper-bright);border-radius:20px;gap:0;margin:0;padding:0;display:grid;overflow:hidden}.fact-table>div{border-bottom:1px solid var(--line);font-size:var(--text-small);grid-template-columns:minmax(96px,30%) minmax(0,1fr);align-items:baseline;gap:6px 24px;padding:13px 20px;display:grid}.fact-table>div:last-child{border-bottom:0}.fact-table dt{color:var(--muted);font-size:var(--text-caption);letter-spacing:.08em;text-transform:uppercase;font-weight:780}.fact-table dd{margin:0;font-weight:620}.profile-layout{grid-template-columns:minmax(0,1fr) 260px;align-items:start;gap:56px;padding-bottom:clamp(56px,9vw,110px);display:grid}.profile-toc{border:1px solid var(--line);background:var(--paper-bright);font-size:var(--text-small);border-radius:20px;gap:8px;padding:22px;display:grid;position:sticky;top:22px}.profile-toc>span{color:var(--muted);font-size:var(--text-caption);letter-spacing:.1em;text-transform:uppercase;font-weight:800}.profile-toc a{color:var(--ink-soft);text-decoration:none}.profile-toc a:hover{color:var(--sea-text);text-decoration:underline}.feature-list{gap:12px;margin:0 0 1.4em;padding:0;list-style:none;display:grid}.feature-list li{border:1px solid var(--line);background:var(--paper-bright);font-size:var(--text-small);border-radius:14px;grid-template-columns:26px minmax(0,1fr);align-items:start;gap:12px;padding:15px 18px;line-height:1.6;display:grid}.feature-list li:before{background:var(--amber-soft);width:26px;height:26px;color:var(--amber-text);content:counter(list-item);font-size:var(--text-caption);border-radius:999px;place-items:center;font-weight:800;display:grid}.feature-list{counter-reset:list-item}.feature-list 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h2{font-family:var(--font-ui);font-size:var(--text-caption);letter-spacing:.13em;text-transform:uppercase;color:var(--muted);margin:0 0 4px;font-weight:800}.footer-nav-groups a{color:var(--ink-soft);font-size:var(--text-small);text-decoration:none}.footer-nav-groups a:hover{color:var(--sea-text);text-decoration:underline}.site-footer small{color:var(--muted);font-size:var(--text-meta)}.home-hero{grid-template-columns:minmax(0,1.15fr) minmax(0,.85fr);align-items:center;gap:52px;padding-block:clamp(56px,9vw,120px);display:grid}.hero-actions{flex-wrap:wrap;align-items:center;gap:18px 26px;margin-top:34px;display:flex}.hero-card{border:1px solid var(--line);background:var(--paper-bright);box-shadow:var(--shadow);border-radius:28px;padding:32px;position:relative;overflow:hidden}.hero-card svg.hero-trilobite{width:100%;height:auto}.hero-card 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0;padding:0;list-style:none;display:grid}.search-results li a{border:1px solid var(--line);background:var(--paper-bright);border-radius:18px;padding:20px 24px;text-decoration:none;display:block}.search-results li a:hover{border-color:var(--sea)}.search-results h2{margin:0 0 4px;font-size:1.16rem}.search-results p{color:var(--muted);font-size:var(--text-small);margin:0}.search-results span{color:var(--amber-text);font-size:var(--text-caption);letter-spacing:.12em;text-transform:uppercase;margin-bottom:8px;font-weight:800;display:inline-block}.not-found{text-align:left;place-content:center;gap:22px;max-width:660px;min-height:74vh;display:grid}.not-found-search{flex-wrap:wrap;gap:12px;display:flex}.not-found-search input{border:1px solid var(--control-border);background:var(--control-bg);color:var(--ink);border-radius:999px;flex:260px;padding:14px 20px}.not-found-links{font-size:var(--text-small);flex-wrap:wrap;gap:10px 20px;font-weight:700;display:flex}@media (width<=1080px){.explorer{grid-template-columns:minmax(0,1fr)}.explorer-filters{position:static}.profile-layout{grid-template-columns:minmax(0,1fr)}.profile-toc{order:-1;position:static}}@media (width<=960px){.desktop-nav,.header-search-link{display:none}.mobile-nav{display:block}.home-hero,.profile-hero,.split-heading,.strip-grid{grid-template-columns:minmax(0,1fr)}.site-header{min-height:72px}}@media (width<=640px){.shell,.shell-narrow{width:calc(100% - 32px)}.fact-table>div{grid-template-columns:minmax(0,1fr)}.prose{font-size:1.02rem}}@media (prefers-reduced-motion:reduce){html{scroll-behavior:auto}*{transition-duration:.01ms!important;animation-duration:.01ms!important}}@media print{.site-header,.site-footer,.profile-toc,.explorer-filters{display:none}}.filter-groups{gap:18px;display:grid}.filter-groups>summary{color:var(--muted);font-size:var(--text-caption);letter-spacing:.1em;text-transform:uppercase;cursor:pointer;font-weight:800}@media (width>=1081px){.filter-groups>summary{display:none}}